The animal rights campaigner has died at the age of 91.

Animal charities and veterinary organisations have paid tribute to animal rights campaigner Jane Goodall, following her death at the age of 91.

Dr Goodall, a primatologist and anthropologist, was considered one of the world’s leading experts on chimpanzees and other great apes, and was responsible for groundbreaking discoveries into primate behaviour.

Her legacy includes a 65-year study of wild chimpanzees living in Gombe, Tanzania, which she began at the age of 26. It was during her time on this project that Dr Goodall gained her understanding of chimpanzee behaviours and her unique bond with the species.

It was also where she made the discovery that chimpanzees used tools, an observation which was said to be ‘redefining what it means to be human’.

However Dr Goodall’s mentor, paleontologist Louis Leakey, knew that her work would only be taken seriously if she had academic qualifications.

Despite her not holding an undergraduate degree, Dr Goodall was able to study for her PhD in Ethology at Newnham College, Cambridge. Her thesis on chimpanzee behaviour in Tanzania was published in 1965.

In 1977 she founded the Jane Goodall Institute, which now has 25 offices operating a number of diverse programmes around the world.

Her later career saw her become a global advocate for animal welfare, species and environmental protection, and for human rights.

Her legacy continues with her ongoing research and conservation programs. She also founded Roots & Shoots, a humanitarian and environmental program for young people of all ages, and published 27 books for adults and children.

Les Ward, chairman of the International Animal Welfare Protection Coalition (IAWPC), said: “Jane was more than a conservationist, she was a beacon of compassion for all life.

“Her support for our campaign gave us courage at a critical moment, and her voice – wise, kind, and unflinching – reminding the world that the animal kingdom deserves respect and that their lives matter. I had the honour of meeting Jane on a number of occasions and each time, I felt that I was in the company of someone truly special.”

Thomas Schultz-Jagow, director of advocacy and prevention at the RSPCA, said: “Dr Jane was truly an inspiration to all at the RSPCA and we share the deep sadness felt around the world following her passing.

“But we also know that her legacy for animals and young people will continue, and we at the RSPCA will continue to share her vision of redefining the relationship between humans and animals around the world."

Elizabeth Mullineaux, senior vice president of the British Veterinary Association, said: "We’re saddened to hear of the passing of Dr Goodall.

"She will be remembered fondly and with admiration by many around the world for her dedication to conservation and protecting our planet. She leaves behind an incredible legacy.”

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